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described by the stationary and progressive waves in fluids, and likewise with the waves of sound. This coincidence would seem to argue an intimate relation between the instrument and the medium on which it is destined to operate— the wing acting in those very curves into which the atmosphere is naturally thrown in the transmission of sound. Can it be that the animate and inanimate world reciprocate, and that animal bodies are made to impress the inanimate in precisely the same manner as the inanimate impress each other? This much seems certain :-The wind communicates to the water similar impulses to those communicated to it by the fish in swimming; and the wing in its vibrations impinges upon the air as an ordinary sound does. The extremities of quadrupeds, moreover, describe waved tracks on the land when walking and running; so that one great law apparently determines the course of the insect in the air, the fish in the water, and the quadruped on the land.

We are, unfortunately, not taught to regard the travelling surfaces and movements of animals as correlated in any way to surrounding media, and, as a consequence, are apt to consider walking as distinct from swimming, and walking and swimming as distinct from flying, than which there can be no greater mistake. Walking, swimming, and flying are in reality only modifications of each other. Walking merges into swimming, and swimming into flying, by insensible gradations. The modifications which result in walking, swimming, and flying are necessitated by the fact that the earth affords a greater amount of support than the water, and the water than the air.

That walking, swimming, and flying represent integral parts of the same problem is proved by the fact that most quadrupeds swim as well as walk, and some even fly; while many marine animals walk as well as swim, and birds and insects walk, swim, and fly indiscriminately. When the land animals, properly so called, are in the habit of taking to the water or the air; or the inhabitants of the water are constantly taking to the land or the air; or the insects and birds which are more peculiarly organized for flight, spend much of their time on the land and in the water; their organs of locomo

tion must possess those peculiarities of structure which charac terize, as a class, those animals which live on the land, in the water, or in the air respectively.

In this we have an explanation of the gossamer wing of the insect,—the curiously modified hand of the bat and bird, -the webbed hands and feet of the Otter, Ornithorhynchus, Seal, and Walrus,-the expanded tail of the Whale, Porpoise, Dugong, and Manatee,—the feet of the Ostrich, Apteryx, and Dodo, exclusively designed for running,—the feet of the Ducks, Gulls, and Petrels, specially adapted for swimming,and the wings and feet of the Penguins, Auks, and Guillemots, especially designed for diving. Other and intermediate modifications occur in the Flying-fish, Flying Lizard, and Flying Squirrel; and some animals, as the Frog, Newt, and several of the aquatic insects (the Ephemera or May-fly for example1) which begin their career by swimming, come ultimately to walk, leap, and even fly.2

Every degree and variety of motion, which is peculiar to the land, and to the water- and air-navigating animals as such, is imitated by others which take to the elements in question secondarily or at intervals.

Of all animal movements, flight is indisputably the finest. It may be regarded as the poetry of motion. The fact that a creature as heavy, bulk for bulk, as many solid substances, can by the unaided movements of its wings urge itself through

1 The Ephemera in the larva and pupa state reside in the water concealed during the day under stones or in horizontal burrows which they form in the banks. Although resembling the perfect insect in several respects, they differ materially in having longer antennæ, in wanting ocelli, and in possessing horn-like mandibles; the abdomen has, moreover, on each side a row of plates, mostly in pairs, which are a kind of false branchiæ, and which are employed not only in respiration, but also as puddles.-Cuvier's Animal Kingdom, p. 576. London, 1840.

2 Kirby and Spence observe that some insects which are not naturally aquatic, do, nevertheless, swim very well if they fall into the water. They instance a kind of grasshopper (Acrydium), which can paddle itself across a stream with great rapidity by the powerful strokes of its hind legs.--(Introduction to Entomology, 5th edit., 1828, p. 360.) Nor should the remarkable discovery by Sir John Lubbock of a swimming insect (Polynema natans), which uses its wings exclusively as fins, be overlooked.-Linn. Traus. vol xxiv. p. 135.

the air with a speed little short of a cannon-ball, fills the mind with wonder. Flight (if I may be allowed the expression) is a more unstable movement than that of walking and swimming; the instability increasing as the medium to be traversed becomes less dense. It, however, does not essentially differ from the other two, and I shall be able to show in the following pages, that the materials and forces employed in flight are literally the same as those employed in walking and swimming. This is an encouraging circumstance as far as artificial flight is concerned, as the same elements and forces employed in constructing locomotives and steamboats may, and probably will at no distant period, be successfully employed in constructing flying machines. Flight is a purely mechanical problem. It is warped in and out with the other animal movements, and forms a link of a great chain of motion which drags its weary length over the land, through the water, and, notwithstanding its weight, through the air. To understand flight, it is necessary to understand walking and swimming, and it is with a view to simplifying our conceptions of this most delightful form of locomotion that the present work is mainly written. The chapters on walking and swimming naturally lead up to those on flying.

In the animal kingdom the movements are adapted either to the land, the water, or the air; these constituting the three great highways of nature. As a result, the instruments by which locomotion is effected are specially modified. This is necessary because of the different densities and the different degrees of resistance furnished by the land, water, and air respectively. On the land the extremities of animals encounter the maximum of resistance, and occasion the minimum of displacement. In the air, the pinions experience the minimum of resistance, and effect the maximum of displacement; the water being intermediate both as regards the degree of resistance offered and the amount of displacement produced. The speed of an animal is determined by its shape, mass, power, and the density of the medium on or in which it moves. It is more difficult to walk on sand or snow than on a macadamized road. In like manner (unless the travelling

surfaces are specially modified), it is more troublesome to swim than to walk, and to fly than to swim. This arises from the displacement produced, and the consequent want of support. The land supplies the fulcrum for the levers formed by the extremities or travelling surfaces of animals with terrestrial habits; the water furnishes the fulcrum for the levers formed by the tail and fins of fishes, sea mammals, etc.; and the air the fulcrum for the levers formed by the wings of insects, bats, and birds. The fulcrum supplied by the land is immovable; that supplied by the water and air movable. The mobility and immobility of the fulcrum constitute the principal difference between walking, swimming, and flying; the travelling surfaces of animals increasing in size as the medium to be traversed becomes less dense and the fulcrum more movable. Thus terrestrial animals have smaller travelling surfaces than amphibia, amphibia than fishes, and fishes than insects, bats, and birds. Another point to be studied in connexion with unyielding and yielding fulcra, is the resistance offered to forward motion. A land animal is supported by the earth, and experiences little resistance from the air through which it moves, unless the speed attained is high. Its principal friction is that occasioned by the contact of its travelling surfaces with the earth. If these are few, the speed is generally great, as in quadrupeds. A fish, or sea mammal, is of nearly the same specific gravity as the water it inhabits; in other words, it is supported with as little or less effort than a land animal. As, however, the fluid in which it moves is more dense than air, the resistance it experiences in forward motion is greater than that experienced by land animals, and by insects, bats, and birds. As a consequence fishes are for the most part elliptical in shape; this being the form calculated to cleave the water with the greatest ease. A flying animal is immensely heavier than the air. The support which it receives, and the resistance experienced by it in forward motion, are reduced to a minimum. Flight, because of the rarity of the air, is infinitely more rapid than either walking, running, or swimming. The flying animal receives support from the air by increasing the size of its travelling surfaces, which act after the manner of twisted

inclined planes or kites. When an insect, a bat, or a bird is launched in space, its weight (from the tendency of all bodies to fall vertically downwards) presses upon the inclined planes or kites formed by the wings in such a manner as to become converted directly into a propelling, and indirectly into a buoying or supporting power. This can be proved by experiment, as I shall show subsequently. But for the share which the weight or mass of the flying creature takes in flight, the protracted journeys of birds of passage would be impossible. Some authorities are of opinion that birds even sleep upon the wing. Certain it is that the albatross, that prince of the feathered tribe, can sail about for a whole hour without once flapping his pinions. This can only be done in virtue of the weight of the bird acting upon the inclined planes or kites formed by the wings as stated.

The weight of the body plays an important part in walking and swimming, as well as in flying. A biped which advances by steps and not by leaps may be said to roll over its extremities,1 the foot of the extremity which happens to be upon the ground for the time forming the centre of a circle, the radius of which is described by the trunk in forward motion. In like manner the foot which is off the ground and swinging forward pendulum fashion in space, may be said to roll or rotate upon the trunk, the head of the femur forming the centre of a circle the radius of which is described by the advancing foot. A double rolling movement is thus established, the body rolling on the extremity the one instant, the extremity rolling on the trunk the next. During these movements the body rises and falls. The double rolling movement is necessary not only to the progression of bipeds, but also to that of quadrupeds. As the body cannot advance without the extremities, so the extremities cannot advance without the body. The double rolling movement is necessary to continuity of motion. If there was only one movement there would be dead points or halts in walking and running, similar to what occur in leaping. The continuity of movement necessary to progression in some bipeds (man for instance) is fur

1 This is also true of quadrupeds. It is the posterior part of the feet which is set down first.

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